1994 Graduated from Kobe University School of Medicine. 2000 Completed the doctoral course in Bioscience, Nara Institute of Science and Technology. 2000 JSPS Research Fellow, Nagoya University. 2003 HFSP Long-term fellow, University of California at San Francisco. 2005 Research fellow, National Center for Geriatrics and Gerontology. 2007 Research fellow, NIPS. 2007 PRESTO Researcher, JST. 2007 Associate Professor, NIPS.
Neuroscience, Biochemistry, Cell Biology