1985 Graduated from Faculty of Medicine, the University of Tokyo. 1989 Completed the doctoral course of Graduate School of Medical Science, the University of Tokyo. 1988-1990 Visiting Scientist, University of Göteborg, Sweden. 1989-1993 Assistant Professor, Institute for Brain Research, the University of Tokyo. 1993-1995 Lecturer & Associate Professor, Gunma University Schoolof Medicine. 1996-2015 Professor, the National Institute for Physiological Sciences. 2015 Professor, Kyoto University Graduate School of Medicine. 2022 Dean, Kyoto University Graduate School of Medicine. 2025 Director General, the National Institute for Physiological Sciences.
