2002 Graduated from Hokkaido University Faculty of Veterinary Medicine. 2004 JSPS Research Fellow (DC2). 2006 Completed the doctoral course in Graduate School of Veterinary Medicine, Hokkaido University, PhD (Veterinary Medicine). 2006 Research Fellow, Research Center for Zoonosis Control, Hokkaido University, 2007 Assistant Professor, Graduate School of Veterinary Medicine, Hokkaido University. 2008 Visiting Research Fellow, Rocky Mountain Laboratories, NIH, USA. 2010, Lecturer, Graduate School of Veterinary Medicine, Hokkaido University. 2018 Assistant Professor, Institute for Genetic Medicine, Hokkaido University. 2021 Lecturer, Institute for Genetic Medicine, Hokkaido University. 2021 Project Associate Professor, NIPS
